g. Hela, 1976 and Lehmann and Myrberg, 2008); i.e. that the thermocline reaches the surface in the upwelling area, bringing cold water from deep layers to the sea surface. This means in practice that our method is only applicable to strong upwelling events taking place in coastal waters. Such common, strong upwelling events,

where a clear drop of SST will take place, could contribute for example, to replenishing the euphotic zone with the nutritional components necessary for biological productivity. Two methods were utilized here to detect and quantify upwelling events. Forskolin For the visual detection method a horizontal grid with longitudinal resolution of 0.5° and latitudinal resolution of 0.25° resulting in a grid box about 28 km2 was overlain on each SST map. As an example Figure 2a shows the SST map for the week 18–25 September 1996 and the overlain grid. It shows that upwelling is occurring along the Polish coast, the Baltic east coast, the west coast of the islands of Saaremaa and Hiiumaa, the Estonian coast of the Gulf of Finland and the Finnish coast of the Bothnian Sea (Figure 2b). For every weekly SST map, upwelling was individually identified and marked in the corresponding box. By doing so, locations within the defined grid and the frequencies of upwelling along the coast of the Baltic Sea could be registered in 443 matrices. For the automatic detection method,

the full resolution of the satellite SST maps was utilized. EGFR inhibitor A simple temperature threshold value was specified. For most parts of the year there exists a latitudinal SST gradient from south to north. Thus, upwelling was detected by calculating the temperature difference for each individual pixel from the zonal mean temperature, for Urease every pixel line. To test the sensitivity of this method with respect to the temperature threshold, two different values (2 °C and 3.5 °C) were specified. For both thresholds erroneous upwelling areas were detected far offshore. Thus, upwelling was only registered if it occurred within a 28 km zone off the coast.

Again, 443 SST maps were scanned and 443 matrices were created but now with a much greater horizontal resolution compared with the visual method. The automatic detection method was also applied to the modelled SST maps, resulting in 3060 matrices showing the location and frequencies of upwelling on the model grid. This method has its limitations if the zonal mean temperature is calculated mainly parallel to the coast such as for the Gulf of Finland, and in spring or autumn when the SST is higher/lower in the coastal area than in the open sea. So we cross-checked upwelling frequencies derived by the automatic method with the results of the visual method. For the wind analysis, the average direction of the different coastal sections was determined from high-resolution bathymetric maps of the Baltic Sea. According to the Ekman theory, winds parallel to the coast are the most effective for causing upwelling.